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The cosmopolitan Crassulaceae comprises about 177 species in the Mediterranean region which, with respect to species richness, is the third centre of diversity within the family. Except for Hylotelephium telephium and Rhodiola roma, the Mediterranean Crassulaceae are endemic to the region or Eurasia. The Crassulaceae originated in Africa, but attained its greaiest diversity tribus Sedge after arrival on the Northern Hemisphere. Most successful is subtribus Sedinae which comprises ihe large, polyphyletic genus Sedum and its derivatives. Subtribus Sedinae comprises two lineages, the Acre-lineage and Leucosedum-lineage, which differ in morphological and molecular characters, but have similar distribution patterns. In Eurasia the Leterosedum-lineage was the most successful, comprising about 75% of all Mediterranean species of Crassulaceae. In addition to some 69 Sedum species it gave rise to the segregate genera eon um, Aichryson, Monanthes, Prometheum, Rosularia, Pistorinia, and Sempervivum (Jovibarba). Although the cen-tres of speciation of these segregate genera as well as the infrageneric taxa of Sedwn are located in different parts of the Mediterranean region there is a distinct parallelism in several evolutionary trends within these to a, e.g. development of the rosulate habit, the polymerous and/or sympetalous flower, and the glandular and often fragrant indumentum.
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