In the swamp forests of Kibale National Park, Uganda, every step elephants take can give rise to a footprint-shaped mini-pond, holding up to 200 liters of water and dozens of invertebrate species. Surveying 30 such prints over a three-day period in 2014. Wolfram Remmers and his colleagues found over 60 species, including beetles, spiders and worms--plus tadpoles. The footprints probably play an important role in allowing these small life forms to spread, as they form a network of connected ponds.
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